Ecological succession begins on bare rock surfaces through pioneering crustose lichens. As lichens decompose, they provide nutrients for mosses to grow. Moss growth traps more soil particles and creates suitable conditions for herbs. Herbaceous plants further weather the rock and add humus. Shrubs later invade and are followed by trees, which constitute the climax forest community. Multiple intermediate seral stages typically evolve over time as the ecosystem advances towards its climax conditions.