Populations grow until reaching the carrying capacity of their environment, limited by factors like food availability. Populations evolve over time through forces like mutation, gene flow, and natural selection. Ecosystems consist of communities of interacting species and their shared physical environment. Over time, ecosystems change through succession as new habitats are colonized by pioneer species and later species replace them. Energy and nutrients cycle through ecosystems via food chains, webs, and biogeochemical cycles, decreasing at each trophic level.