PHYLUM ANNELIDA (Segmented worms)
- 1. M ANNEL IDA Submitted by: Ninna Pauline Caraan ClariceTeresa Guinto IV - Nitrogen
- 2. Introduction ▪ Annelida is a group commonly referred to as segmented worms, and they are found worldwide from the deepest marine sediments to the soils in our city parks and yards. Through most of the 20th century Annelida was split into three major groups; Polychaeta, Oligochaeta (earthworms etc.) and Hirudinea (leeches).
- 3. ▪ Earthworms and leeches are the familiar annelids for most people, but polychaetes comprise the bulk of the diversity of Annelida and are found in nearly every marine habitat, from intertidal algal mats downwards.
- 4. There are even pelagic polychaetes that swim or drift, preying on other plankton, and a few groups occurring in fresh water and moist terrestrial surroundings. Around 9000 species of polychaetes are currently recognized with several thousand more names in synonymy, and the overall systematics of the group remains unstable (Rouse and Pleijel, 2001). Introduction
- 7. CLASSES ▪ Class Polychaeta ▪ Class Oligochaeta ▪ Class Hirudinea
- 8. POLYCHAETA: Bristle Worms ▪ The vast majority of the more than 8,000 known species of polychaete worms are marine; some, however, are found in fresh or brackish water.They are abundant from the intertidal zone to depths of over 16,405 ft (5,000 m). The Sea Mouse The BomberWormsDiopatra
- 9. ▪ The polychaetes, so named because of the numerous setae (chaetae) they bear, range in length from less than 1/8 in. to more than 9 ft (2 mm to 3 m), but most are from 2 to 4 in. (5–10 cm) long.Their colors are often brilliant, and some species are iridescent.The class has usually been divided on the basis of mode of existence into two groups, the errantia and the sedentaria. CLASS POLYCHAETA: Bristle Worms Fireworms, Eurythoe spp (A), C - Spirorbis, 'fanworm' or 'feather duster' worm (B)
- 10. Errant Polychaeta ▪ Errant polychaetes include actively crawling or swimming forms which may, however, also spend time in burrows or crevices, or under rocks on the seashore. A familiar errant polychaete is the clamworm, Nereis, widely used as bait.
- 11. Sedentary Polychaetes ▪ Sedentary polychaetes are usually adapted to living permanently in tubes or burrows; some attach themselves to rocks or piers. Many sedentary polychaetes, like the lugworm, Arenicola, live in burrows in sand or mud. The majority, however, are tube builders.
- 12. Polychaete Anatomy The structure of the digestive tract of polychaetes is variable, reflecting the diversity of feeding types. Respiration is entirely through the body wall in some polychaetes, and partially so in most.
- 14. Above: a 3D model of a pair of segments of a typical errant polychaete, as viewed from the anterior end. Polychaete Anatomy
- 15. ▪ The tentacles of feather-duster worms are used for respiratory exchange as well as for feeding. A polychaete may have a single pair of excretory tubes or a pair in each segment. Sedentary polychaetes have various modifications to insure that wastes will be deposited near the mouth of the tube or burrow, where they are washed away. Many species have thin-walled extensions of the body surface, i.e., gills, used for gas exchange; most commonly the gills are extensions of the parapodia. Polychaete Anatomy
- 16. POLYCHAETE REPRODUCTION ▪ Most polychaetes reproduce sexually, and the sexes are separate. Sex cells develop from masses of tissue in the metameres and leave by way of tubules or by rupture of the body wall. In most cases fertilization of the eggs by sperm occurs externally in seawater and results in the formation of free-swimming larvae.Variations include internal fertilization, laying of egg masses that are attached to objects with mucus, and brooding of developing eggs in the worm's body.
- 18. CLASS OLIGOCHAETA ▪ This class includes about 3,500 species of earthworms and freshwater worms.The members of the class range in length from about 1/32 in. to 10 ft (0.5 mm–3 m), but most are comparable to the polychaetes in size. Oligochaetes occur in a variety of habitats throughout the world. Most are burrowers in the soil, but the class also includes worms that inhabit wells, marshes, and swamps. Other species live under rocks on the seashore, in the leaves of tropical trees and vines, on the surface of glaciers, or on the gills of freshwater crayfish.
- 20. ▪ Like the polychaetes, oligochaetes have bodies divided into segments. However, they lack parapodia and, with a few exceptions, have relatively few and inconspicuous setae.The setae are usually arranged in four bundles on each segment; those of aquatic forms are longer than those of land forms.The setae of an earthworm may be felt as a roughness if one rubs a finger along its side. OLIGOCHAETA ANATOMY
- 22. ▪ Oligochaetes are less varied in their external form than the polychaetes, but are much more numerous. ▪ As many as 4,000 oligochaetes have been counted in 1 square meter of lake bottom, and about 9,000 in 1 square meter of meadow soil. ▪ In almost all oligochaetes, the head is a simple cone-shaped structure without sensory appendages. ▪ Light is detected by photoreceptor cells in the skin, usually concentrated toward the front of the animal. OLIGOCHAETA ANATOMY
- 23. OLIGOCHAETE DIGESTION ▪ The mouth, located under the head, leads to a relatively simple, straight digestive tract consisting of a pharynx, an esophagus, and an intestine, terminating in an anal opening.
- 24. ▪ Terrestrial oligochaetes tunnel through the ground, swallowing soil as they go.The digestive tract of such a worm is specially modified for this rough diet.Typically it has a thin-walled storage area, or crop, and a muscular gizzard for grinding the soil to remove the organic matter that is the actual food of the worm. OLIGOCHAETE DIGESTION
- 25. OLIGOCHAETE CIRCULATION AND RESPIRATION ▪ The circulatory system is that typical of the annelids and has many contractile vessels, or hearts. Although a few aquatic forms have gills for respiration, most oligochaetes lack such specialized structures and use the capillaries of their body walls for respiratory exchange. Oxygen dissolved in the soil water diffuses through the moist epidermis of the worm.
- 26. ▪ Earthworms have a variety of sensory cells.Touch cells, or mechanoreceptors, contain tactile hairs, which, when stimulated, trigger a nerve impulse.The earthworm also has light-sensitive cells. Such cells are more abundant in its anterior and posterior segment, the parts of its body most likely to be outside of the burrow. One of the earthworm's most sensitive cells are those that detect moisture.These cells are located on the first few segments.The animal also appears to have taste cells. In the laboratory, worms can be shown to select celery in preference to cabbage leaves. OLIGOCHAETE NERVOUS SYSTEM
- 27. OLIGOCHAETE REPRODUCTION All oligochaetes are hermaphroditic, and nearly all cross-fertilize by copulation. Male and female reproductive organs are located in separate segments. The copulating pair exchange sperm, which are stored in the body of the recipient worm until its eggs are mature.
- 29. ▪ This class includes the 500 species of leeches, flattened, predacious or parasitic annelids equipped with suckers used for creeping. Leeches range in length from about 1/2 in. to 8 in. (1 cm–20 cm); most are under 2 in. (5 cm) long.They are commonly black, brown, green, or red, and may have stripes or spots. CLASS HIRUDINEA
- 30. ▪ Leeches are primarily freshwater annelids, but some live in the ocean and some in moist soil or vegetation.The majority of leeches are predators on small invertebrates; most swallow their prey whole, but some suck the soft parts from their victims. Some leeches are parasites rather than predators, and suck the body fluids of their victims without killing them.The distinction is not sharp, as many predatory leeches take blood meals on occasion.
- 31. HIRUDINEA Distinguishing characteristics ▪ Distinguishing characteristics - Leeches have suckers at both their oral and anal ends (Fig. B).They are usually flat so they appear thin from the side view (Fig. C).They are relatively smooth, lacking setae like other annelids (Fig. D). Leeches have variable coloration, ranging from light to dark, spotty to striped (Fig. E).
- 33. HIRUDINEA Locomotion ▪ More complex muscle system, "inchworm" movement via suckers
- 34. HIRUDINEA Reproduction ▪ Hermaphrodites, with separate testis and ovaries.The sexual organs, and the ducts that lead to and from them are situated in the anterior (front) part of the animal, normally between segments 7 and 15.
- 35. ▪ A courtship display is seen among some leeches at the time of mating.The fertilized eggs are deposited in a cocoon, secreted by the clitellum; the cocoon is buried in mud or affixed to submerged objects.The young emerge as small copies of the adults. HIRUDINEA Reproduction Cont.