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VTS 150 Animal Nutrition Fall 2010 Instructor:  Beth Alden, DVM
Course Objectives Understanding of different digestive systems Learn the nutritional needs of small companion animals, “pocket pets,” large animals and exotic pets Learn about nutritional diseases for several species Learn how to calculate an animal’s energy needs Know how to evaluate a pet food label and ingredients Learn common toxicities for dogs and cats
Resources for Class Textbook : Nutrition for Veterinary  Technicians and Nurses ,  Ann Wortinger , Blackwell Publishing, 2007
Resources on Library Reserve Small Animal Clinical Nutrition 4 th  Edition,   Hand,Thatcher,  Remillard, Roudebush,   The Mark Morris Institute, 2000
What’s Required of You: 2 Unit Exams Three (3) Student Projects Final Exam
Digestive System Classifications
Digestive Classifications Two Methods of  Digestive System Classification Food Source Stomach Type
Herbivores:  Plant eaters Carnivores: Meat Eaters Omnivores: Plant and Meat Eaters
Herbivores Plant material makes up diet Rough vegetation, nitrogen source, minerals & water Difficult to digest Plant cell wall is hard to break down  Energy from the plant is tied up in cellulose that animal cannot digest on its own The animal has developed a large complex gut to digest this material Microbial action in gut is necessary to break down the cellulose and release its energy Little of the food’s energy is absorbed by host
Carnivores Animal material (flesh, muscle, etc.) makes up diet Food more easily digested than in herbivores The animal has a very simple “gut “ Most of food’s energy is absorbed by host Mechanical breakdown of food is less involved
Types of Digestive Systems Four compartment stomach: Reticulum, Rumen, Omasum & Abomasum (cows, sheep, goats) One “true” stomach (dogs, cats, pigs, horses) Ruminant Monogastric
Monogastric Animals
The Monogastric Stomach Five sections: Cardia Fundus Body Antrum Pylorus Source: University of California at Davis
The Monogastric Stomach 1. Cardia Area immediately  surrounding the opening from the esophagus into the stomach  Muscular tone prevents reflux of stomach contents into the esophagus CARDIA Source: University of California at Davis
The Monogastric Stomach 2. Fundus Located below the cardia Blind pouch that distends as food is swallowed Source: University of California at Davis FUNDUS CARDIA
The Monogastric Stomach 3. Body Distends to receive food Rich with gastric glands Parietal Cells : Hydrochloric acid (HCl) Chief Cells :  Pepsinogen Mucous Cells : Mucus BODY Source: University of California at Davis FUNDUS CARDIA BODY
The Monogastric Stomach 4. Antrum Grinds up food and regulates HCl Also contains glands G Cells:  Gastrin Mucous Cells :  Mucus BODY Source: University of California at Davis FUNDUS CARDIA BODY ANTRUM
The Monogastric Stomach 5. Pylorus Muscular ring (sphincter)  Regulates movement of  chyme  from stomach into duodenum Helps prevent backflow of duodenal contents BODY Source: University of California at Davis FUNDUS CARDIA BODY ANTRUM PYLORUS DUODENUM
Ruminant Animals
The Ruminant Animal Chews and swallows its food then regurgitates “cud” to chew on it more before swallowing again Has a “ prestomach ” configuration that is adapted to the herbivore diet Four compartment stomach
The Ruminant Stomach Four Compartments: Reticulum Rumen Omasum Abomasum Source:  University of California at Davis head tail
The Ruminant Stomach 1. The Reticulum Smallest, most cranial compartment Separated from rumen by the  ruminoreticular fold Muscular wall is continuous with the rumen; contract in coordination RETICULUM Source:  University of California at Davis head tail
The Ruminant Stomach 1. The Reticulum Honeycombed inside to increase absorptive surface “ Hardware disease”- wires or nails swallowed by animal puncture wall of reticulum RETICULUM Inside of Reticulum Source:  Colorado State University
The Ruminant Stomach 2. The Rumen Large fermentative vat (40 - 50 gallon capacity) Processes plant material into usable energy Lined with  “Papillae” Made up of series of muscular   “ Pillars ” RUMEN Source:  University of California at Davis head tail
The Ruminant Stomach © University of Bristol, 1988 Pillar Papillae Papillae
The Ruminant Stomach 2. The Rumen During contractions, pillars close off certain sacs of the rumen which allows mixing of ruminal contents Mixing of contents essential for fermentative function of rumen RUMEN
The Ruminant Stomach What reticuloruminal contractions do : Allow “cud” to be regurgitated up the esophagus, where it’s re-chewed and  re-swallowed Allow  “eructation”  - built-up CO 2   or methane gas to be expelled from rumen.  Gasses are forced into the reticulum and  up the esophagus RUMEN
The Ruminant Stomach Fermentative digestion begins in the rumen Bacterial and protozoal enzymes break down foodstuffs Carbohydrate Metabolism : Cellulase  enzymes digest cellulose (plant cell layer) and transform complex carbohydrates into simple sugars Simple sugars aren’t made available to host (like they are in monogastric animals), instead are absorbed into the microbes and converted to  Volatile Fatty Acids (VFAs) RUMEN
The Ruminant Stomach Carbohydrate Metabolism,  continued Volatile fatty acids are by-products of anaerobic fermentation by bacteria in rumen Some VFAs absorbed into liver then converted to glucose, which is used by the animal’s cells Other VFAs are used to produce adipose tissue and milk fat RUMEN
The Ruminant Stomach Protein Metabolism: Rumen microbial enzymes attack proteins  (just like carbohydrates) Proteases  reduce long proteins to  amino acids  and short  peptides  (short chains of amino acids) Peptides are either incorporated into the protein structure of the microbes OR converted to ammonia (NH 3 + ) and VFAs RUMEN
The Ruminant Stomach Protein Metabolism,  continued Liver secretes  urea*  into the rumen, which provides additional nitrogen for the microbes Microbes are then flushed from the reticulorumen to the omasum, abomasum and intestines; digested and serve as source of protein to the animal * Urea is sometimes added to poor-quality feed as a way of increasing nitrogen value RUMEN
The Ruminant Stomach Other Notes about the Rumen Provides B vitamins; Vitamin K Delicate balance: amount  of food, growth of microbes  and by-products Changes in diet severely affect production of gas, VFAs and ammonia, causing fermentation process to completely stop RUMEN
The  Ruminant Stomach 3.  Omasum Muscular organ located off the reticulum Ingesta moves into omasum from reticulorumen Prevents large particles from leaving rumen and entering abomasum OMASUM Source:  University of California at Davis head tail
The Ruminant Stomach 3. Omasum Primary functions Break down food particles further and move ingesta to abomasum Absorb any excess VFAs  Remove bicarbonate ions from ingesta (to avoid altering acid pH of abomasum) OMASUM
The Ruminant Stomach 4. Abomasum “ True stomach” of ruminant Functions similar to  monogastric stomach ABOMASUM Source:  University of California at Davis head tail
Young Ruminant Digestive Tract Functions as a monogastric system No fermentative digestion  (rumen & reticulum are non-functional) Reticular groove  or  Esophageal groove  allows suckled milk to go to omasum Suckling is important – helps groove to function  Bucket–fed calves spill milk into rumen & reticulum
Young Ruminant Digestive Tract Abomasum is largest of 4 stomachs for the  first few weeks Rumen & reticulum development    rate dependent on diet: Grain & Hay fed – develop at 3 weeks Milk fed – develop at 3 months
Chronology of Digestive System GI tract extends from mouth to the anus and performs different functions at different parts Chronology of Digestion : Prehension Mechanical grinding down of food Chemical digestion of food Absorption of nutrients and water Elimination of waste material
Chronology of Digestion 1.  Prehension Grasping food with teeth or lips
Chronology of Digestion : 2. Mechanical Grinding Down of Food Carnivore Teeth Pointed to facilitate holding and tearing of food ©  Clinical Textbook for Veterinary Technicians 6 th  ed .; McCurnin, Bassert
Chronology of Digestion: 2. Mechanical Grinding Down of Food Herbivore Teeth Flat, occlusal surface for grinding
Chronology of Digestion: 2. Mechanical Grinding Down of Food Incisors  – Teeth in the front for holding and tearing Canines  – Pointed teeth located at corners for tearing and shredding Premolars  – Located just before the molars and are used for grinding in all species Molars  – Used for grinding
Chronology of Digestion: 2. Mechanical Grinding Down of Food Ruminant Teeth No upper incisors or upper canine teeth Dental Pad Thick connective tissue Diastema
Chronology of Digestion: 2. Mechanical Grinding Down of Food Positional Location of Teeth - Terminology Maxilla Upper Arcade Mandible Lower Arcade Lingual Inner side of lower arcade teeth  that face tongue Labial Outer surface of upper and lower arcade teeth (faces lips) Palatal Inner side of upper arcade teeth (faces the palate) Buccal Outer side of teeth in back of mouth  (faces the cheeks)
Chronology of Digestion: 3.  Chemical Digestion of Food Saliva mixes with food during chewing  Three bilateral pairs of salivary glands in the mouth  ( parotid, mandibular, lingual ) What saliva does: softens, moistens, shapes & lubricates food aids in taste acts as a buffer provides digestive enzymes
Chronology of Digestion 3.  Chemical Digestion of Food Digestive enzymes and buffers in saliva Amylase :   in omnivore saliva – not carnivore  breaks down amylose  (sugar component of starch) Lipase: breaks down lipids Bicarbonate and Phosphate buffers: in cow saliva; neutralizes acids in rumen & maintain normal rumen pH  ( up to 25-30 gallons of saliva a day)
Chronology of Digestion 3.  Chemical Digestion of Food Epiglottis   Cartilage projection that prevents food from entering trachea Esophagus   - transports food to stomach Peristalsis: smooth muscle contractions to propel food
Chronology of Digestion 3.  Chemical Digestion of Food Stomach   Stores food  Continues enzymatic breakdown of food (pepsin, gastrin, hydrochloric acid) Mechanical breakdown of food (mixing, grinding and contractions that move food) Ruminants:  each compartment has different function
Chronology of Digestion 3.  Chemical Digestion of Food Liver Secretes bile acids to help  with digestion of fat Pancreas Secretes enzymes into small intestine for breaking down nutrients: Protease  for proteins Amylase  for carbohydrates Lipase  for fats/lipids Also secretes bicarbonate to  neutralize stomach acid
Chronology of Digestion 4.  Absorption of Nutrients & Water Small Intestine Duodenum, Jejunum, Ileum Inner & sub mucosal layer;  Muscular layer Villi & Microvilli increase surface area in intestine Peristaltic waves help move contents through intestines
Chronology of Digestion 4.  Absorption of Nutrients & Water Small Intestine No clear demarcation  between 3 segments All 3 segments perform  peristalsis, absorb nutrients & H 2 O Duodenum:   first portion of sm. intestine Jejunum:   majority of sm. intestine Ileum:   where sm. intestine enters colon
Chronology of Digestion 4.  Absorption of Nutrients & Water Villi Millions of cylindrical, fingerlike projections in intestinal wall Provide large surface area for absorbing nutrients “ Crypts”  surround villi and replenish cells Microvilli  “brush border” Found on Villi; contain digestive enzymes Transmissible Gastroenteritis (TGE) in pigs Parvo Virus in dogs attack the villus  Animal cannot absorb  nutrients from  intestinal tract
Chronology of Digestion 4.  Absorption of Nutrients & Water Small Intestine Absorbs electrolytes  (Na, Cl, K, etc.); water and vitamins across  intestinal wall Absorbs carbohydrates, fats, proteins after they’re chemically digested via enzymes
Chronology of Digestion 4.  Absorption of Nutrients & Water Nutrient digestion in the small intestine Carbohydrates:   digested by amylase secreted from pancreas Protein: digested by proteases secreted by pancreas Fat: digested by bile acids from liver  (helps emulsify fat to prevent glob  formation); further broken down  by lipase secreted from pancreas
Chronology of Digestion 5.  Elimination of Waste Material Large Intestine:  Cecum & Colon Recover fluid and electrolytes Store feces until elimination Some microbial action Differences between species, dependent on diet
Chronology of Digestion 5.  Elimination of Waste Material *Horses are different Colon Cecum    “blind sac” Carnivores Simple, tubular, contracts to move feces through Poorly developed Herbivores* Large bacterial population (microbes)  for fermentation More developed, larger than carnivore
Chronology of Digestion 5. Elimination of Waste Material Colon & Cecum comprise the  “Hindgut” 4 sections: cecum, ventral colon, dorsal colon, small colon More highly developed than small intestine Has greater capacity for fermentation Unique digestion path;  colonic impaction is most common form of colic in horses
Chronology of Digestion 5. Elimination of Waste Material Rectum Terminal portion of colon Contains mucus-secreting glands Sensory receptors detect stretching/distension and triggers defecation Anus Internal & External sphincters allow controlled passage  of feces
Next Week… Session 2:  Basic Nutrients

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Session 1 Orientation And Animal Classifications

  • 1. VTS 150 Animal Nutrition Fall 2010 Instructor: Beth Alden, DVM
  • 2. Course Objectives Understanding of different digestive systems Learn the nutritional needs of small companion animals, “pocket pets,” large animals and exotic pets Learn about nutritional diseases for several species Learn how to calculate an animal’s energy needs Know how to evaluate a pet food label and ingredients Learn common toxicities for dogs and cats
  • 3. Resources for Class Textbook : Nutrition for Veterinary Technicians and Nurses , Ann Wortinger , Blackwell Publishing, 2007
  • 4. Resources on Library Reserve Small Animal Clinical Nutrition 4 th Edition, Hand,Thatcher, Remillard, Roudebush, The Mark Morris Institute, 2000
  • 5. What’s Required of You: 2 Unit Exams Three (3) Student Projects Final Exam
  • 7. Digestive Classifications Two Methods of Digestive System Classification Food Source Stomach Type
  • 8. Herbivores: Plant eaters Carnivores: Meat Eaters Omnivores: Plant and Meat Eaters
  • 9. Herbivores Plant material makes up diet Rough vegetation, nitrogen source, minerals & water Difficult to digest Plant cell wall is hard to break down Energy from the plant is tied up in cellulose that animal cannot digest on its own The animal has developed a large complex gut to digest this material Microbial action in gut is necessary to break down the cellulose and release its energy Little of the food’s energy is absorbed by host
  • 10. Carnivores Animal material (flesh, muscle, etc.) makes up diet Food more easily digested than in herbivores The animal has a very simple “gut “ Most of food’s energy is absorbed by host Mechanical breakdown of food is less involved
  • 11. Types of Digestive Systems Four compartment stomach: Reticulum, Rumen, Omasum & Abomasum (cows, sheep, goats) One “true” stomach (dogs, cats, pigs, horses) Ruminant Monogastric
  • 13. The Monogastric Stomach Five sections: Cardia Fundus Body Antrum Pylorus Source: University of California at Davis
  • 14. The Monogastric Stomach 1. Cardia Area immediately surrounding the opening from the esophagus into the stomach Muscular tone prevents reflux of stomach contents into the esophagus CARDIA Source: University of California at Davis
  • 15. The Monogastric Stomach 2. Fundus Located below the cardia Blind pouch that distends as food is swallowed Source: University of California at Davis FUNDUS CARDIA
  • 16. The Monogastric Stomach 3. Body Distends to receive food Rich with gastric glands Parietal Cells : Hydrochloric acid (HCl) Chief Cells : Pepsinogen Mucous Cells : Mucus BODY Source: University of California at Davis FUNDUS CARDIA BODY
  • 17. The Monogastric Stomach 4. Antrum Grinds up food and regulates HCl Also contains glands G Cells: Gastrin Mucous Cells : Mucus BODY Source: University of California at Davis FUNDUS CARDIA BODY ANTRUM
  • 18. The Monogastric Stomach 5. Pylorus Muscular ring (sphincter) Regulates movement of chyme from stomach into duodenum Helps prevent backflow of duodenal contents BODY Source: University of California at Davis FUNDUS CARDIA BODY ANTRUM PYLORUS DUODENUM
  • 20. The Ruminant Animal Chews and swallows its food then regurgitates “cud” to chew on it more before swallowing again Has a “ prestomach ” configuration that is adapted to the herbivore diet Four compartment stomach
  • 21. The Ruminant Stomach Four Compartments: Reticulum Rumen Omasum Abomasum Source: University of California at Davis head tail
  • 22. The Ruminant Stomach 1. The Reticulum Smallest, most cranial compartment Separated from rumen by the ruminoreticular fold Muscular wall is continuous with the rumen; contract in coordination RETICULUM Source: University of California at Davis head tail
  • 23. The Ruminant Stomach 1. The Reticulum Honeycombed inside to increase absorptive surface “ Hardware disease”- wires or nails swallowed by animal puncture wall of reticulum RETICULUM Inside of Reticulum Source: Colorado State University
  • 24. The Ruminant Stomach 2. The Rumen Large fermentative vat (40 - 50 gallon capacity) Processes plant material into usable energy Lined with “Papillae” Made up of series of muscular “ Pillars ” RUMEN Source: University of California at Davis head tail
  • 25. The Ruminant Stomach © University of Bristol, 1988 Pillar Papillae Papillae
  • 26. The Ruminant Stomach 2. The Rumen During contractions, pillars close off certain sacs of the rumen which allows mixing of ruminal contents Mixing of contents essential for fermentative function of rumen RUMEN
  • 27. The Ruminant Stomach What reticuloruminal contractions do : Allow “cud” to be regurgitated up the esophagus, where it’s re-chewed and re-swallowed Allow “eructation” - built-up CO 2 or methane gas to be expelled from rumen. Gasses are forced into the reticulum and up the esophagus RUMEN
  • 28. The Ruminant Stomach Fermentative digestion begins in the rumen Bacterial and protozoal enzymes break down foodstuffs Carbohydrate Metabolism : Cellulase enzymes digest cellulose (plant cell layer) and transform complex carbohydrates into simple sugars Simple sugars aren’t made available to host (like they are in monogastric animals), instead are absorbed into the microbes and converted to Volatile Fatty Acids (VFAs) RUMEN
  • 29. The Ruminant Stomach Carbohydrate Metabolism, continued Volatile fatty acids are by-products of anaerobic fermentation by bacteria in rumen Some VFAs absorbed into liver then converted to glucose, which is used by the animal’s cells Other VFAs are used to produce adipose tissue and milk fat RUMEN
  • 30. The Ruminant Stomach Protein Metabolism: Rumen microbial enzymes attack proteins (just like carbohydrates) Proteases reduce long proteins to amino acids and short peptides (short chains of amino acids) Peptides are either incorporated into the protein structure of the microbes OR converted to ammonia (NH 3 + ) and VFAs RUMEN
  • 31. The Ruminant Stomach Protein Metabolism, continued Liver secretes urea* into the rumen, which provides additional nitrogen for the microbes Microbes are then flushed from the reticulorumen to the omasum, abomasum and intestines; digested and serve as source of protein to the animal * Urea is sometimes added to poor-quality feed as a way of increasing nitrogen value RUMEN
  • 32. The Ruminant Stomach Other Notes about the Rumen Provides B vitamins; Vitamin K Delicate balance: amount of food, growth of microbes and by-products Changes in diet severely affect production of gas, VFAs and ammonia, causing fermentation process to completely stop RUMEN
  • 33. The Ruminant Stomach 3. Omasum Muscular organ located off the reticulum Ingesta moves into omasum from reticulorumen Prevents large particles from leaving rumen and entering abomasum OMASUM Source: University of California at Davis head tail
  • 34. The Ruminant Stomach 3. Omasum Primary functions Break down food particles further and move ingesta to abomasum Absorb any excess VFAs Remove bicarbonate ions from ingesta (to avoid altering acid pH of abomasum) OMASUM
  • 35. The Ruminant Stomach 4. Abomasum “ True stomach” of ruminant Functions similar to monogastric stomach ABOMASUM Source: University of California at Davis head tail
  • 36. Young Ruminant Digestive Tract Functions as a monogastric system No fermentative digestion (rumen & reticulum are non-functional) Reticular groove or Esophageal groove allows suckled milk to go to omasum Suckling is important – helps groove to function Bucket–fed calves spill milk into rumen & reticulum
  • 37. Young Ruminant Digestive Tract Abomasum is largest of 4 stomachs for the first few weeks Rumen & reticulum development rate dependent on diet: Grain & Hay fed – develop at 3 weeks Milk fed – develop at 3 months
  • 38. Chronology of Digestive System GI tract extends from mouth to the anus and performs different functions at different parts Chronology of Digestion : Prehension Mechanical grinding down of food Chemical digestion of food Absorption of nutrients and water Elimination of waste material
  • 39. Chronology of Digestion 1. Prehension Grasping food with teeth or lips
  • 40. Chronology of Digestion : 2. Mechanical Grinding Down of Food Carnivore Teeth Pointed to facilitate holding and tearing of food © Clinical Textbook for Veterinary Technicians 6 th ed .; McCurnin, Bassert
  • 41. Chronology of Digestion: 2. Mechanical Grinding Down of Food Herbivore Teeth Flat, occlusal surface for grinding
  • 42. Chronology of Digestion: 2. Mechanical Grinding Down of Food Incisors – Teeth in the front for holding and tearing Canines – Pointed teeth located at corners for tearing and shredding Premolars – Located just before the molars and are used for grinding in all species Molars – Used for grinding
  • 43. Chronology of Digestion: 2. Mechanical Grinding Down of Food Ruminant Teeth No upper incisors or upper canine teeth Dental Pad Thick connective tissue Diastema
  • 44. Chronology of Digestion: 2. Mechanical Grinding Down of Food Positional Location of Teeth - Terminology Maxilla Upper Arcade Mandible Lower Arcade Lingual Inner side of lower arcade teeth that face tongue Labial Outer surface of upper and lower arcade teeth (faces lips) Palatal Inner side of upper arcade teeth (faces the palate) Buccal Outer side of teeth in back of mouth (faces the cheeks)
  • 45. Chronology of Digestion: 3. Chemical Digestion of Food Saliva mixes with food during chewing Three bilateral pairs of salivary glands in the mouth ( parotid, mandibular, lingual ) What saliva does: softens, moistens, shapes & lubricates food aids in taste acts as a buffer provides digestive enzymes
  • 46. Chronology of Digestion 3. Chemical Digestion of Food Digestive enzymes and buffers in saliva Amylase : in omnivore saliva – not carnivore breaks down amylose (sugar component of starch) Lipase: breaks down lipids Bicarbonate and Phosphate buffers: in cow saliva; neutralizes acids in rumen & maintain normal rumen pH ( up to 25-30 gallons of saliva a day)
  • 47. Chronology of Digestion 3. Chemical Digestion of Food Epiglottis Cartilage projection that prevents food from entering trachea Esophagus - transports food to stomach Peristalsis: smooth muscle contractions to propel food
  • 48. Chronology of Digestion 3. Chemical Digestion of Food Stomach Stores food Continues enzymatic breakdown of food (pepsin, gastrin, hydrochloric acid) Mechanical breakdown of food (mixing, grinding and contractions that move food) Ruminants: each compartment has different function
  • 49. Chronology of Digestion 3. Chemical Digestion of Food Liver Secretes bile acids to help with digestion of fat Pancreas Secretes enzymes into small intestine for breaking down nutrients: Protease for proteins Amylase for carbohydrates Lipase for fats/lipids Also secretes bicarbonate to neutralize stomach acid
  • 50. Chronology of Digestion 4. Absorption of Nutrients & Water Small Intestine Duodenum, Jejunum, Ileum Inner & sub mucosal layer; Muscular layer Villi & Microvilli increase surface area in intestine Peristaltic waves help move contents through intestines
  • 51. Chronology of Digestion 4. Absorption of Nutrients & Water Small Intestine No clear demarcation between 3 segments All 3 segments perform peristalsis, absorb nutrients & H 2 O Duodenum: first portion of sm. intestine Jejunum: majority of sm. intestine Ileum: where sm. intestine enters colon
  • 52. Chronology of Digestion 4. Absorption of Nutrients & Water Villi Millions of cylindrical, fingerlike projections in intestinal wall Provide large surface area for absorbing nutrients “ Crypts” surround villi and replenish cells Microvilli “brush border” Found on Villi; contain digestive enzymes Transmissible Gastroenteritis (TGE) in pigs Parvo Virus in dogs attack the villus Animal cannot absorb nutrients from intestinal tract
  • 53. Chronology of Digestion 4. Absorption of Nutrients & Water Small Intestine Absorbs electrolytes (Na, Cl, K, etc.); water and vitamins across intestinal wall Absorbs carbohydrates, fats, proteins after they’re chemically digested via enzymes
  • 54. Chronology of Digestion 4. Absorption of Nutrients & Water Nutrient digestion in the small intestine Carbohydrates: digested by amylase secreted from pancreas Protein: digested by proteases secreted by pancreas Fat: digested by bile acids from liver (helps emulsify fat to prevent glob formation); further broken down by lipase secreted from pancreas
  • 55. Chronology of Digestion 5. Elimination of Waste Material Large Intestine: Cecum & Colon Recover fluid and electrolytes Store feces until elimination Some microbial action Differences between species, dependent on diet
  • 56. Chronology of Digestion 5. Elimination of Waste Material *Horses are different Colon Cecum “blind sac” Carnivores Simple, tubular, contracts to move feces through Poorly developed Herbivores* Large bacterial population (microbes) for fermentation More developed, larger than carnivore
  • 57. Chronology of Digestion 5. Elimination of Waste Material Colon & Cecum comprise the “Hindgut” 4 sections: cecum, ventral colon, dorsal colon, small colon More highly developed than small intestine Has greater capacity for fermentation Unique digestion path; colonic impaction is most common form of colic in horses
  • 58. Chronology of Digestion 5. Elimination of Waste Material Rectum Terminal portion of colon Contains mucus-secreting glands Sensory receptors detect stretching/distension and triggers defecation Anus Internal & External sphincters allow controlled passage of feces
  • 59. Next Week… Session 2: Basic Nutrients