Puccinia is a genus of fungi that includes approximately 5000 species, some of which are devastating pathogens of wheat and other cereals. Puccinia graminis causes stem rust in wheat, P. recondita causes brown leaf rust, and P. striformis causes yellow stripe rust. These rusts have polymorphic life cycles with multiple spore stages. Symptoms vary by species but generally include blisters or pustules that erupt and release spores, which can be brick red in stem rust and cause stems and leaves to blacken. Brown leaf rust first appears on upper leaves while yellow stripe rust primarily affects leaves. In barberry shrubs, rust symptoms appear as yellow to orange spots on leaves.