This document discusses several spatial models used to simulate energy flow and nutrient dynamics in ecosystems. It describes the CENTURY model, which simulates grassland ecosystems, and how it has been combined with GIS data to model spatial patterns in carbon and nitrogen storage. It also discusses the FOREST-BGC model for simulating forest carbon, nitrogen, and water cycles over broad landscapes, and the FIRE-BGC model which incorporates the effects of fire disturbances into FOREST-BGC. The document emphasizes that spatial variations in environmental factors can produce substantial variation in ecosystem processes, and that models need to operate at appropriate spatial scales to develop predictive relationships.