The Dynamic Language Runtime (DLR) allows .NET languages to execute dynamically like dynamic languages such as Python and Ruby. The DLR builds on the CLR to provide services for dynamic languages, including dynamic types, dynamic method dispatch, and code generation. It uses expression trees to represent language semantics and call site caching for faster dynamic method dispatch. The DLR enables interoperability between statically and dynamically typed languages. It includes types in the System.Dynamic namespace and language binders that allow it to interact with objects from other languages like JavaScript, IronRuby, IronPython, and COM.